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Berbers are primarily not African ?
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[QUOTE]Originally posted by the lioness,: [QB] TAFORALT MAN IN SAHARA : SAHARAN EXTENSION OF MAGHREBIAN 1988 ^^^ this article refers to a possible later extension of Iberomaurusians of Tafroalt, Morocco into Hassi-el-Abiod in Mali. This would fit the edn of wet phase transition driving this partcular robust cromagnid population south Skeletal remains in both of these areas are temperate proportioned not tropical _________________________________________________ [b]Mitochondrial Haplogroup H1 in North Africa: An Early Holocene Arrival from Iberia[/b] 2010 Claudio Ottoni equal contributor, Giuseppina Primativo equal contributor, Baharak Hooshiar Kashani,Alessandro Achilli, Cristina Martínez-Labarga, Gianfranco Biondi, Antonio Torroni, Olga Rickards mail Published: October 21, 2010 Abstract [b]The Tuareg of the Fezzan region (Libya) are characterized by an extremely high frequency (61%) of haplogroup H1, a mitochondrial DNA (mtDNA) haplogroup that is common in all Western European populations.[/b] To define how and when H1 spread from Europe to North Africa up to the Central Sahara, in Fezzan, we investigated the complete mitochondrial genomes of eleven Libyan Tuareg belonging to H1. Coalescence time estimates suggest an arrival of the European H1 mtDNAs at about 8,000–9,000 years ago, while phylogenetic analyses reveal three novel H1 branches, termed H1v, H1w and H1x, which appear to be specific for North African populations, but whose frequencies can be extremely different even in relatively close Tuareg villages. Overall, these findings support the scenario of an arrival of haplogroup H1 in North Africa from Iberia at the beginning of the Holocene, as a consequence of the improvement in climate conditions after the Younger Dryas cold snap, followed by in situ formation of local H1 sub-haplogroups. This process of autochthonous differentiation continues in the Libyan Tuareg who, probably due to isolation and recent founder events, are characterized by village-specific maternal mtDNA lineages. [IMG]http://www.plosone.org/article/fetchObject.action?uri=info:doi/10.1371/journal.pone.0013378.g001&representation=PNG_M[/IMG] Entrance into the Levant paved the way to the dispersal of modern humans both north-westward into Europe and south-westward into Northern Africa [10], [11]. During the Last Glacial Maximum (LGM), approximately between 26.5 and 19 kya [12] ice sheets largely covered large portions of North America and Europe. In warmer regions of the world, the climate was cooler and drier and deserts spread over large regions, particularly in Northern Africa, Middle East and Central Asia [13], [14]. Accordingly, during the LGM, humans concentrated in refugial areas of southwestern Europe, in the Balkans and Levant, and on the east European plains [9], [15], [16]. The subsequent Břlling warming, around 15 kya, triggered re-expansion processes which led to the resettlement of Central and Northern Europe. Genetic signatures of these expansions are evident in mtDNA genealogies, for instance haplogroups H1, H3 and V contributed to the gradual re-peopling of Europe from the Franco-Cantabrian refuge in the postglacial [17], [18]. Similarly, though to a lesser extent, H5*(xH5a), H20 and H21 may be associated to a postglacial population expansion phase in the Caucasus area [19]. Although restricted to the Mediterranean coast, an expansion took place also from the Italian peninsula northward, as attested to by the haplogroup U5b3 [20]. Evidence of trans-Mediterranean contacts between Northern Africa and Western Europe has been assessed at the level of different genetic markers (e.g. [21], [22], [23], [24]). With regards to the mtDNA, the high incidence of H1 and H3 in Northwest Africa, together with some other West European lineages (i.e. V and U5b), reveals a possible link with the postglacial expansion from the Iberian Peninsula, which not only directed north-eastward into the European continent [17], [18], [25], but also southward, beyond the Strait of Gibraltar, into North Africa [26], [27]. So, besides the ‘autochthonous’ South-Saharan component, the maternal pool of Northern Africa appears to be characterized by at least two other major components: (i) a Levantine contribution (i.e. haplogroups U6 and M1, [11]), associated with the return to Africa around 45 kya, and (ii) a more recent West European input associated with the postglacial expansion. Within the West-European component in North Africa, H1 is the most represented haplogroup with frequencies ranging from 21% in some Tunisian Berber groups to 1% in Egypt [28]. Recently, an extremely high incidence of H1 (61%) has been reported in a Tuareg population from the Central Sahara, in Libya [29]. Tuareg are a semi-nomadic pastoralist people of Northwest Africa, who speak a Berber language. MtDNA analyses performed on the Libyan Tuareg have highlighted their genetic relatedness with some Berber groups and other North African populations, mainly resulting from the sharing of a common West-Eurasian component. A high degree of homogeneity in the Libyan H1 lineages was observed, suggesting that the high frequency of H1 in the Tuareg may be the result of genetic drift and recent founder events. To better define the nature and extent of H1 variation in the Tuareg from Libya we have now determined the complete sequence of eleven of their mtDNAs belonging to H1. The comparison of these H1 sequences with those already available from Europe and North Africa provides new clues on how and when H1 spread in Northern Africa up to the Central Sahara. Overall, the results of this study support the hypothesis that most of the West Eurasian maternal contribution detectable in Northwest African populations is likely linked to prehistoric (i.e. the post-glacial expansion from the Iberian Peninsula) rather than more recent historic events [26], [27], [37]. Furthermore, the data presented confirm that the analysis of complete mtDNA sequences represents a valuable tool to reveal not only the spatial patterns beneath large-scale colonization events, but also those of smaller-scale dispersals which may have contributed to the origin of modern populations. In this regard, additional efforts in the full mtDNA analyses of nomad Northern African populations might resolve the debate concerning their origin and their mutual relationship. [/QB][/QUOTE]
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