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[QUOTE]Originally posted by Clyde Winters: [QB] [IMG]http://i44.tinypic.com/293bwup.jpg[/IMG] . R1 originated in Africa and spread into Eurasia. As a result haplogroup R is very diverse in Africa.There is a great diversity of the macrohaplogroup R in Africa (See Figure 1). Ychromosome R is characterized by M207/V45. The V45 mutation is found among African populations ( Cruciani et al ,2010). ISOGG 2010 Y-DNA haplogroup tree makes it clear that V45 is phylogenetically equivalent to M207.The most common R haplogroup in Africa is R1 (M173). The predominant haplogroup is R1b (Berniell-Lee et al,2009; Coia et al, 2005; Winters, 2010b; Wood et al, 2009). Cruciani et al (2010) discovered new R1b mutations including V7, V8, V45, V69, and V88. [IMG]http://olmec98.net/r1.jpg[/IMG] I specifically stated the frequency of R1 among African populations throughout my 2011 paper. [QUOTE] Y-chromosome R1 is found throghout Africa. The pristine form of R1-M173 is only found in Africa (Coia et al, 2005; Cruciani et al, 2002, 2010). The age of y chromosome R is 27ky. Most researchers believe that R(M173) is 18.5 ky.There is a great diversity of the macrohaplogroup R in Africa (See Figure 1). Ychromosome R is characterized by M207/V45. The V45 mutation is found among African populations ( Cruciani et al ,2010). ISOGG 2010 Y-DNA haplogroup tree makes it clear that V45 is phylogenetically equivalent to M207.The most common R haplogroup in Africa is R1 (M173). The predominant haplogroup is R1b (Berniell-Lee et al,2009; Coia et al, 2005; Winters, 2010b; Wood et al, 2009). Cruciani et al (2010) discovered new R1b mutations including V7, V8, V45, V69, and V88. Geography appears to play a significant role in the distribution of haplogroup R in Africa. Cruciani et al (2010) has renamed the R*- M173 (R P-25) in Africa V88. The TMRCA of V88 was 9200-5600 kya (Cruciani et al, 2010). Y-chromosome V88 (R1b1a) has its highest frequency among Chadic speakers, while the carriers of V88 among Niger-Congo speakers (predominately Bantu people) range between 2-66% ( Cruciani et al, 2010; Bernielle-Lee et al, 2009). Haplogroup V88 includes the mutations M18, V35 and V7. Cruciani et al (2010) revealed that R-V88 is also carried by Eurasians including the distinctive mutations M18, V35 and V7.[b] R1b1-P25 is found in Western Eurasia. Haplogroup R1b1* is found in Africa at various frequencies. BerniellLee et al (2009) found in their study that 5.2% carried Rb1*. The frequency of R1b1* among the Bantu ranged from 2-20. The bearers of R1b1* among the Pygmy populations ranged from 1-25% (Berniell-Lee et al, 2009). The frequency of R1b1 among Guinea-Bissau populations was 12% (Carvalho et al,2010).[/b] [/QUOTE][IMG]http://olmec98.net/carriers249.jpg[/IMG] In relation to R-M269 in Africa I wrote: [QUOTE] Around 0.1 of Sub Saharan Africans carry R1b1b2. Wood et al (2009) found that Khoisan (2.2%) and Niger-Congo (0.4%) speakers carried the R-M269 y-chromosome. The Khoisan also carry RM343 (R1b) and M 198 (R1a1) (Naidoo et al., 2010) the archaeological and linguistic data indicate the successful colonization of Asia by SubSaharan Africans from Nubia 5-4kya (Winters, 2007,2008, 2010c). The archaeological evidence makes it clear that around 4kya intercultural style artifacts connected Africa and Eurasia (Winters, 2007,2010c). [/QUOTE]As a result, I did not attempt to decieve anyone about the frequency of R1 in Africa as the author implies. In fact recent research on y-haplogroups in Africa suggest that R1-M269 is also widespread in Africa. [IMG]http://olmec98.net/m269.png[/IMG] Above is a figure from Gonzalez et al. The Gonzalez et al article found that 10 out of 19 subjects in the study carried R1b1-P25 or M269. This is highly significant because it indicates that 53% of the R1 carriers in this study were M269, this finding is further proof of the widespread nature of this so-called Eurasian genes in Africa among populations that have not mated with Europeans. The R1 haplogroup probably originated in Africa. . . [IMG]http://olmec98.net/phylogenyR1.png[/IMG] . The phylogeography of R1 in Africa makes it clear that this y-chromosome is spread globally across Africa and includes the genetic structure of diverse African populations including Berber, Chadic, Cushitic, Khoisan,Pygmy, Niger-Congo, Nilo-Saharan and Semitic speaking African populations (Berniell-Lee et al, 2009; Cruciani et al, 2010; Wood et al, 2009). The fact that Dravidians carry the R haplogroup illustrate the recent introduction of R y-chromosome to Eurasia. Abu-Amero et al (20009) reveal the fact that Dravidians carry the R haplogroups illustrate the recent introduction of Ry-chromosomes to Eurasia. The frequency of haplotype M173 in Eurasia is as follows: Anatolia 0.19%, Iran 2.67%, Iraq 0.49% Oman 1.0%, Pakistan 0.57% and Oman 1.0% . This contrast sharply with the widespread distribution of R1 in Africa that ranges between 7- 95% in various parts of Africa, especially Cameroon (Coia et al, 2005). Coia et al (2005) has revealed that no maternal Eurasian lineages have been found among Sub-Saharan Africans with a R1- M173 profile. Haplogroup V88 has the greatest frequency in Africa. It is predominately carried by Chadic speakers, ranges between 2-60% among Central African Niger-Congo speakers (Cruciani et al, 2010). Researchers have found that the TMRCA of V88 was 9200-5600 kya (Cruciani et al, 2010). [IMG]http://olmec98.net/r1.jpg[/IMG] The vast majority of Africans belong to the y-chromosome E macrohaplogroup. Phylogenetically haplogroup R1b is mainly found in West Africa and the Sahel. In this region the frequency of R-M173 can range between 85-100% among some Niger Congo speakers in Cameroon (Cruciani et al, 2010). The paternal record of M173 on the African continent illustrates a greater distribution of this y-chromosome among varied African populations than, in Asia. [IMG]http://olmec98.net/Carriers2492.jpg[/IMG] The greatest diversity of R1b in Africa is highly suggestive of an Africa origin for this male lineage because it is not isolated to just one part of Africa. Archaeological (Lal, 1963), genetic (Winters, 2008;2010a), placenames (Balakrishnan, 2005) and linguistic data group (Aravanan,1979,1980; Upadhyaya, 1976,1979; Winters 1985a,1985b, 1989) linking Africans and Dravidian support the recent demic diffusion of SubSaharan Africans and gene flow from Africa to Eurasia. An early colonization of Eurasia 4kya by Sub-Saharan Africans and Dravidian carriers of R1-M173 is the best scenario to explain the high frequency and widespread geographical distribution of this y-chromosome on the African continent (Winters, 2010c). Given the greatest diversity of R1- M173, this is the most parsimonious model explaining the frequency of R-M173 in Africa. Africans carry haplogroup R1a. In India the Dravidian people carry the R1a haplogroup The Dravidian people of India originally lived in Middle Africa and belonged to the Proto-Saharan Civilization. The Proto-Saharan civilization was situated in the Proto-Sahara, which includes Cameroon. . [IMG]http://olmec98.net/blee.png[/IMG] . In Cameroon we find carriers of R1a. In addition to carriers of R1a in Cameroon; the [b]Dravidian languages are still spoken today in Cameroon see:[/b] https://www.youtube.com/watch?v=vWyAYGlFZjkhttps://www.youtube.com/watch?v=vWyAYGlFZjk In conclusion, the R macrohaplogroup probably originated in Africa. In my paper POSSIBLE AFRICAN ORIGIN OF Y-CHROMOSOME R1-M173 , I argue that the P clade originated in Africa because 1) the age of R-V88 and 2) the widespread nature of R1 in Africa. Researchers have found that the TMRCA of V88 was 9200-5600 kya (Cruciani et al, 2010). Eurasians carry the M269 (R1b1b2) mutation. The subclades of R1b1b2 include Rh1b1b2g (U106) (TMRCA 8.3kya) and R1b1b2h (U152) (TMRCA 7.4kya). The most recent common ancestor for R1b1b2 is probably 8kya (Balaresque et al, 2010). In Africa we find R-M269 and V88. Clearly, R-V88 is older than R-M269 there is no evidence of archaeological evidence of a back migration or haplogroup R into Africa, but there is evidence of the migration of the Kushites and Proto-Saharans into Eurasia from Middle Africa. This supports the proposition the R haplogroups originated in Africa, not Eurasia. [/QB][/QUOTE]
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